Publications

Thesis

• Espregueira Themudo, G. (2005) Study of the Marbled Newt (Triturus (m.) marmoratus and T. (m.) pygmaeus) hybrid zone by genetic markers. MSc. thesis, University of Porto, Portugal.
• Espregueira Themudo, G. (2010) Newts in time and space: the evolutionary history of Triturus newts at different temporal and spatial scales. Leiden University, The Netherlands.

Book chapters

• Espregueira Themudo, G. and J. W. Arntzen 2008 Triturus marmoratus in: Atlas dos anfíbios e répteis de Portugal (eds. Armando Loureiro, Nuno Ferrand, Miguel Carretero).

Papers in peer-reviewed journals

1. Wielstra, B., Espregueira Themudo, G., Güçlü, Ö., Olgun, K., Poyarkov, N.A., Arntzen, J.W., 2010. Cryptic crested newt diversity at the Eurasian transition: The mitochondrial DNA phylogeography of Near Eastern Triturus newts. Molecular Phylogenetics and Evolution 56: 888-896. (Abstract)
2. Espregueira Themudo, G., B. Wielstra and J. W. Arntzen. (2009). Multiple nuclear and mitochondrial genes resolve the branching order of a rapid radiation of crested newts (Triturus, Salamandridae). Molecular Phylogenetics and Evolution 52: 321-328. (Abstract) (Erratum)
3. Espregueira Themudo, G., W. Babik, and J. W. Arntzen. (2009). A combination of techniques proves useful in the development of nuclear markers in the newt genus Triturus. Molecular Ecology Resources 52: 321-328. (Abstract)
4. Arntzen, J. W., and G. Espregueira Themudo. 2008. Environmental parameters that determine species geographical range limits as a matter of time and space. Journal of Biogeography 35: 1177-1186. (Abstract)
5. Arntzen, J. W., G. Espregueira Themudo, and B. Wielstra (2007) The phylogeny of crested newts (Triturus cristatus superspecies): nuclear and mitochondrial genetic characters suggest a hard polytomy, in line with the paleogeography of the centre of origin. Contributions to Zoology 76:261-278. (Full text) (PDF)
6. Espregueira Themudo, G., and J. W. Arntzen (2007) Newts under siege: range expansion of Triturus pygmaeus isolates populations of its sister species. Diversity and Distributions 13:580-586. (Abstract)
7. Espregueira Themudo, G., and J. W. Arntzen (2007) Molecular identification of marbled newts and a justification of the Triturus marmoratus and T. pygmaeus species status. Herpetological Journal 17:24-30. (Abstract) (PDF)